FOOD: The Behavior Shaper

I’ve written this posting to clear up the difference between food-conditioning and simple acclimation — there seems to be confusion about these terms. 

This coyote pictured here has been listlessly hanging around, within five feet of a path in a park, where he dozes and waits for food to be tossed to him. Food is tossed to him off and on, so he is being rewarded for his efforts.  He has a family he could be with — a pup, a yearling and a mate — but food trumps that for this coyote. He should be hunting — but then again, why would he do that when food will just come his way if he simply lies here? In fact, I have not seen him hunt in a while.

There’s a person who feels he is “protecting” the coyote by letting people know he is not dangerous: “Look, I can go right up to him and he does nothing,”  he tells people multiple times, daily. I tried convincing him that his constant approaching the coyote is not helping matters. This guy also feels the coyote needs to be fed: “He’s hungry, right? or he wouldn’t be there begging for food.” Other people approach to look or photograph him with their iPhones, getting as close as 5-10 feet away: few people carry a good zoom lens which would allow them to keep their distance. And further: they then post the coyote’s location on their social media which draws in even more people to approach or feed and iPhotograph. The feeding incidents take a mere second: it’s hard to catch beforehand even if you are standing right there constantly, so the “no feeding” ordinance is hard to enforce.

I’ve been here educating, but I can’t be here all the time, so I’ve been soliciting as many people as possible to be ambassadors to help others in the area understand that feeding by humans and friendliness — which encourages coyotes to view us as potential feeders — are actually “faux amis”: they are robbing the coyote of his independence and survival skills, and encouraging him to lie around within 5 feet of heavy human pedestrian traffic all day. It’s heart-wrenching to watch if you know coyotes.

Some people have even asked me, “What’s wrong with that, after all, he’s not hurting anyone.” But others are more in-tuned and ask if he is sick, or even dead when he’s dozing off. A handful of people have admitted to me that they had been feeding the coyote regularly — they hadn’t known better — but now they do: they thanked me for the clear signs. The signs I recently put out seem to be yielding some results.

Contrary to what many people have been led to believe, the problem here is not caused by the coyote’s having become acclimated to humans. I know lots of coyotes who have become acclimated to our presence without ending up in our midst or as “problems”. In fact, coyotes throughout the city, in any urban area, are all acclimated to humans by definition: they get used to us because we amicably share the same environment, including in the parks. Be that as it may, almost all remain wary and keep their distance: coyotes don’t just up and start mingling with us simply because they’re in the habit of seeing us or no longer see us as fearsome. Why would they — what would be the draw? Nor is there any “progression” in this acclimatization behavior whereby they eventually come ever closer, and then even become assertive or even threatening towards humans. Yet some people promote this as a truth, using the word “habituation”. It’s a concept causing people to fear the presence coyotes unnecessarily. These people are actually confounding “acclimation” with “food-conditioning”. The two are not the same and have to be kept apart.

“Food conditioning”, when it occurs, on the other hand, especially over time, indeed becomes a problem, and that is what is going on here with this coyote. This coyote’s behavior was not caused by simple acclimation to human presence. The rest of his family does not behave as he does. It was caused by the consistent and persistent proffering of food by friendly humans, so that he now associates humans as a friendly food source. Also keep in mind that every coyote is different, so innate personality plays a role.



Words and their meanings. Exact word meanings are important when talking about such an emotionally charged subject as coyotes, where everyone has a strong pre-conceived opinion. Without using exact language you cannot convey what is really going on or how to deal with it, and this seems to be the case where the meaning of the word “habituation” which is supposed to mean “the diminishing of a physiological or emotional response to a frequently repeated stimulus”, has been expanded to include food-conditioned behaviors: wouldn’t that then be the “increasing of a physiological or emotional response”?  This confounding, then, attributes incorrect causes to certain behaviors. I’ll avoid the word in order to avoid tapping into anyone’s pre-conceived misunderstanding of the term. We need to understand these as two separate phenomenon: “food-conditioning” vs. simple adaptation to humans. I’ll use the word “acclimation” instead.

“Acclimation” is defined as the “the process or result of becoming accustomed to something new.” In this case it means simple “accommodation” and nothing more: the definition is congruent with the italicized definition given above.  Its effect has been noted in all animals including us. So, for instance, by living in the city, we humans learn to ignore and even screen out noises so that we need not waste energy worrying or reacting to something that isn’t going to harm us: these non-threatening sounds include sirens, a blown-out tire, fire-works, or even a rock band in the park.  Acclimation does not cause us to increase our reaction to those non-dangerous things we become accustomed to, it diminishes our need to react.

This is also true of coyotes. When coyotes become used to humans by adapting to our habitual presence — accommodating us — they ignore us because they realize that we are not a danger, that we are simply part of the environment that’s out there. They do not come towards us or beg for food or become aggressive towards us just because they have become used to us. Think about it: why would they? Getting to know humans and our behavior as we go about our normal and separate lives — without trying to intimidate or scare them all the time — actually creates calmer and, yes, LESS reactive coyotes! But scare them all the time and they’re bound to start showing their teeth self-protectively. Walkaboutlou has noted that if you treat coyotes harshly, they’ll become hard coyotes.

Watch the process as it develops to know what is going on: I have been watching specifically this for over 12 years — for coyote reactions over long periods of time from birth to people and our behaviors [dogs and pets are a different issue which I will cover later].

Wariness and keeping distant are actually built into coyote behavior naturally as you can see by the aversive behavior of all youngsters. But this can be trained out of them by humans: food is this behavior shaper. This coyote here is hanging around unnaturally close to human activity: this was not caused by his becoming acclimated to us. What led to this behavior is humans breaching the natural divide by INTERACTING with him first and foremost through friendly feeding. This, then, coupled with befriending, attempting to communicate, approaching, and even prolonged mutual visual contact exacerbated the problem by making all humans potential feeders. These, interactive behaviors by humans, and not simply human presence, are what alter the behavior of coyotes so that they may hang around close to human activity and even follow people in an attempt to get more food: for them, it’s an easier thing to do than hunting. Coyotes are opportunistic and towards that end are constantly pushing their boundaries to their advantage: if it is advantageous for them, they will change their behaviors.

And BTW, I have never seen feeding lead to aggressiveness. In fact the feeding that I’ve observed over time — and it’s always very friendly feeding — results in very docile, meek, and almost tame coyotes who hang around listlessly waiting for food to be tossed their way. They become nuisances more than anything else, and the situation becomes circular and perpetual. Most importantly, this situation could lead to more negative consequences in that this “proximity” could provide opportunities for these animals to grab a kid’s sandwich or even react to a hyperactive small child. My wildlife animal behaviorist contact says that “feeding changes the relationship between a wild animal and humans, putting them on a more equal footing with us, which, if the animal were to become desperate enough it might, potentially, ‘demand’ food from a human. This is not something that is a regular occurrence, but it has happened.” By feeding we are training the animal — shaping the animal’s behavior (talk to any dog owner to find out how food is used to train an animal) to hang around, which could possibly lead to demanding or intrusive behavior. Food is the behavior shaper. Friendliness abets the process.

IN SUM, ALL of the URBAN coyotes that I know are acclimated, and this is due to the urban situation and by definition: they become used to us because we are there –we are ever-present in the parks we share with them. Nevertheless, they naturally keep their distance and only occasionally cross paths with us. They learn to ignore us because we are not a danger to them. We are simply a part of the environment “out there.” This should not be a problem.

But SOME coyotes have been encouraged by people beyond acclimation, to INTERACT on some level with us and become absorbed into our world. Again, every coyote is different, so innate personality will also play a role here. THIS interaction then, is what is unhealthy for everyone: it breaks down the natural safety barriers that were innately in place. It is occurring more frequently due to a pendulum swing from too much fear towards coyotes, to too much love, primarily through feeding, compounded with befriending, interacting with, approaching,. . . . etc.

People need to understand that they are hurting the coyote by interacting — they are shaping the coyote behavior away from its natural state.. Please, always walk away from a coyote, not for your own safety necessarily, though for that too, but for the well-being of the coyotes. Understanding this process is helping many people change their too-friendly behaviors towards coyotes. However, when this education is ignored, maybe it needs to be backed up by enforcement with fines.

Coyotes, too, have attempted to initiate interactions with some dogs as we walk them — it’s a way they use for finding out about these dogs who they see as “intruders” in “their” territories. Coyotes and dogs generally do not like each other, and small pets, of course, can be vulnerable as prey. I’ll get into this in another posting, but it’s important to prevent engagement by simply walking the other way, away from a coyote. If a coyote has approached your dog too closely as you are trying to move away from it, this is when you’ll need to react more pro-actively with anger and intimidation. More on this soon.

Note 1: One of the rationales that has been tossed at me is that feeding coyotes will keep them from grabbing pets. I read where a neighborhood in Los Angeles put out dog food which apparently cut down on disappearing cats. But in fact, you may just be encouraging the coyote to hang around closer to where s/he CAN indeed grab a pet. Even in this case, you would still need to leash your pet to keep it safe especially from chasing the coyote, so why not just start here in the first place and work on keeping away from coyotes?

Note 2: I hope you noticed that this coyote’s ears are hanging low — almost “floppy ears”. I’ve noticed its persistence in fed coyotes. It has been noted by a Russian scientist that this trait grows, and eventually is inherited, as wild dogs, specifically foxes, become tamer. See the famous red fox study about this.

Mountain Lion in San Francisco!

https://youtu.be/UjCp1BXSEOU
Mountain lions in San Francisco? Well, they don’t really live here, but we have had about one a year wander through the city in the last three years. Here’s one that was caught on a security/trap camera last night close to Lake Merced, at the very southern end of the city. If they don’t leave quickly and they appear lost, our Animal Care and Control, along with the California Department of Fish and Wildlife, will tranquilize the animal and remove it both for its own good and that of the public. These animals could pose a potential threat. The last one that was taken out of the city two years ago was a young female weighing 92 pounds. That’s about my size: I wouldn’t want to tangle with one. Coyotes, on the other hand, weigh a mere 30 to 35 pounds and are not a threat to humans. Nevertheless, we need to keep our distance from them, less for our own safety than for the wellbeing of the coyote.

Addendum: I’d like to post here a couple of other animals that one rarely sees in San Francisco, but who passed by my night cameras. A gray fox, and a red fox, and then a video with raccoon, opossum, a hawk, dogs, and a skunk scares off coyotes.  :)

End of Summer Ranch Observations, by Walkaboutlou

Here are some amazing end-of-the-summer observations I wanted to share. There’s so much information here, lots of detail, and incredible insight, beautifully woven together into a letter. Enjoy and learn! Janet

Hi Janet,

Lou here. Summer is ending and I’m piecing together local coyote snippets and news and ranch situations. All told, very consistent with local human behaviors.

On the ranches where no coyote are hunted, (and livestock are cattle) everything seems very “stable” to minimal. Small litters of 2-5. Predictable vocalizations. The usual subtle background living Coyotes seem to enjoy. The scat in these areas is full of plum and apricot seeds, deer hair, tons of blackberries, and overwhelmingly rodents.

Overall, of course each coyote is a fluid and distinctive individual, subject to rapid change and stages.

But if my summer scouting had a theme, it would be the contrast of Coyotes behavior even in similar regions.

For example, non hunted coyote in cattle ranches (4000 acres or more) seem to develop small, stable packs and territory. The food and ecosystem are abundant in large ranches. If the cattle can range, grass grows leaving vast regions of insects, and rodents. The pups learn early to forage on grasshoppers, mice. Very predictable quiet patterns. Often seen in distance in diurnal behaviors. By Fall, usually 2-3 pups remain. (accidents and natural predators curb litter survival) Pups seem to want to hang with pack a year or 2. Also, prey is scavenged until gone. A deer dying from being hit by car (running off to die in brush) or fawns harvested are eaten and visited until gone. Nothing is wasted.

The contrast again in ranches that hunt coyote hard is almost shocking. I have determined large, sheep operations are very challenging for Coyotes to coexist peacefully. If it’s large, LGD can only be in so many places. Also, large herds of sheep graze the land intensely. The cropped grass becomes a giant short lawn, unsuitable habitat for rodents, insects etc..if sheep are grazing long, you’ll notice hardly any sounds of crickets etc…and blackberry bushes are cut by ranchers because sheep get entangled. So the lack of forage, food and cover changes the setting. Add to this intense human hunting. Very intense. The coyote often become nocturnal. The closely cropped land and human hunters do not favor open, relaxed foraging. There are minimal rodents. So the coyote tend to hole up all day and hunt far and wide very hard at night with time as a factor. And pressure. Another complication-large herds of sheep especially isolated always have old, sick, hurt or dead. Or a scattering of lambs in all directions. The coyote scavenge dead sheep, or prey on lambs. Their pups are weaned on sheep. The smell becomes embedded as food-and a cycle is created.

Other reactions caused by human pressure-non hunted coyote females pick a mate around 2nd year. Hunted female coyote often pair bond as yearlings. So daughter’s breed earlier and with larger litters in answer to hunting pressure.

The social ramifications are evident. Many ranchers will hunt and leave a coyote as a magnet for surviving pack members to investigate, becoming targets themselves. Only this doesn’t work long. Hunted Coyotes learn to truly leave the dead behind. Some mothers will not check out a deceased pup or mate. This detachment of survival to me is amazing but sad too.

Also, such hard living Coyotes show other behaviors. They quickly, hurriedly hunt. And more readily raid any livestock or pets that opportunities give. They often do not return to a carcass after one feed. They’ve learned hunters, greyhounds or snares are sometimes waiting.

Pups scatter and really practice independence by Fall. The long puppyhood of stable packs is absent in hunted coyote.

All in all, stable coyote packs and hunted coyote are vastly different. And unfortunately, the unpredictability of hunted coyote makes them unwelcome even among stable packs. They really are different. And bring behaviors that can influence others.

I wish I could just make everyone leave coyote alone. They would still be amazing. But we would see and learn so much more without ignorance or outright war. Coyote are definetly mirrors of the local humans. If I want to know about people’s culture/lifestyle/knowledge or lack of, the local dogs and coyote will inform me.

Keep Studying and Coexisting.

Lou🐾


Hi Janet-I did forget to add one element to my summer coyote scouting.

This pic off internet sums it up well.

In areas where packs of coyote live more or less normally, you’ll find more or less the usual range in size and color of coyote. Especially in West. However, where coyote are hard hunted and scattered year round, you will find some that obviously have more then coyote genetics. This goes in hand with younger females (yearlings) breeding and also lower coyote numbers. If they are hit hard locally, surviving coyote have no hesitation breeding with dogs, especially free roaming ranch dogs (often kelpie/cattle dog/collie types).

This, in turn, can create more variety in local subspecies of coyote-and no doubt affects some. Larger size or bolder demeanor are often traits of 1st generation crosses. They tend to be absolved back into wild populations. But are another aspect of hard hunted coyote.
In laymans words-if you take away a coyote’s mate and think she’s beat, she’ll just recruit your dog as her next husband. And the pups won’t be Lassie. Either way, coyote will turn the dice of man’s efforts into a win.

Lou🐾

Night Vision Snippets: Peeks at Pup Feedings

Here, I’ve consolidated several very short video clips from over a month ago which I caught on a trail camera: pup feedings. You’ll see a mother coyote regurgitating food for a very hungry and excited pup who can’t wait to eat. Then you’ll see the same youngster happily carrying a small rodent off to a quiet place where she can enjoy it. Thirdly, you’ll see Dad on his way to deliver food to his brood at 3am in the morning: you have to wonder how many trips he must make to feed four young mouths. Next, a youngster runs excitedly after Mom and yanks away the prey which she has brought home for the pup. Finally, a pup, now several weeks older than before, carries something large which I can’t really make out in the infrared light. It’s either prey that has been given to her by a parent, or a toy which she has found and is treating as prey.

NOT Coyotes

Every now and then I post something that has nothing at all to do with coyotes, and this posting is one of those: Surfing dogs.

More

Coyote Family Playtime for a 3-Month Old Singleton Pup


This tiny family responds to sirens!

She’s an “only pup” — she has no litter mates. An “only pup” is known as a “singleton” pup. But she is not an “only child” because she has an older brother: a yearling born the year before. He was part of a litter of five, and is the only youngster from that litter to remain part of the family. That yearling plays with the pup, as do Mom and Dad, as you’ll see in the video.

Nighttime is when coyote families engage in most of their family activities: the whole family plays together on and off — when adults aren’t off hunting — during the length of the evening. And then they rest or sleep in different locations during the daytime.

The video above is a composite from one of my rarer daylight captures of family play. Note that, after the intense and fun play session above, the “adults”, trickle off, one at a time, in the end leaving the pup alone for the rest of the day. At night, too, they leave her for long extended periods of time when they go off hunting. She knows she must stay home and keep hidden.

After watching them leave, the pup wanders sadly, slowly, and unenthusiastically back — you can tell this by the lack of energy in her pace — to her hiding place. And that’s how the days go by as she is growing up.

Scout Continues On The Run

Scout disappeared from my radar about six weeks ago, after a short return to her old territory at the end of June from where she left abruptly after seeing that her vanquisher was still there. It appears that this little coyote will never give up hope of returning. . . . some day, some day.

For those who do not know her story, she was viciously driven out of the territory she had owned for three years by another coyote, Wired, in February. Sadly, humans probably had a hand in the outcome of the territorial battle.

July 20th. Photo credit: WMontgrain

A couple of weeks ago she resurfaced in a park across the city. Some of the photos showed her looking well and healthy, but others showed her looking very thin, even factoring in that all coyotes look scrawny at this time of year because it’s the end of their shedding season: there is nothing left of their 3″ thick and fluffy winter coats.

She is an interloper without a territory, and interlopers are not allowed to stay long on claimed territories. The new park has long belonged to another entrenched coyote family. Scout was there for only about ten days, when a neighbor saw the territorial fight on Sunday, July 21st at dusk that seems to have ended her stay there. She has not been seen there since this happened: she is again on the run, leading the life of a fugitive.

So Scout is continuing to experience hard times. I try to console myself be remembering that coyotes are resilient and are made to deal with adversity. We all hope she is okay.

I continue to see Wired very sporadically in the territory she took over from Scout, but few other people have ever seen her there — she is a much more elusive critter. She makes regular long exploratory trips to all corners of the city: her latest excursion took her to 19th Avenue, close to Stern Grove. After these wanderings, Wired always returns to the territory she won from Scout where she continues to be irregularly, but often enough to keep Scout away and homeless. UPDATE 8/8: According to her radio-collar emissions (as per the ecologist at the Presidio) she has spent her recent time at the Presidio and Lands End, and been all the way down to Brisbane.

Trap Cameras: Dad Patrols / Pup Explores

Today, I’m posting a video sequence of a coyote on his patrolling rounds, and of a youngster investigating the area, captured remotely on a trap camera. Pups are 3 months old now and their curiosity is insatiable, of course! But this posting is more about trap cameras which aren’t as benign as most people think.


Use of trap cams and what I’ve seen. I’ve used them extremely sparingly in the past. Out of respect for what coyotes want and so as to observe natural rather than fearful or protective behavior, I stay away from denning areas: rather, I’ve used field cameras to count pups after they start exploring well away from their dens. And I’ve used them more recently for tracking an interloper coyote who was driven from her territory by another coyote.

There IS a difference between trap-cameras and human hand-held cameras, but not necessarily in the way people imagine: I’m referring to the intrusiveness factor. In some ways they may be less interfering than human presence, but in many ways they are more so.

Some of the reactions of coyotes to my and other trap cameras as seen in images from those cameras led to my doing a mini-study on trap-cameras. The outcome: I found that these devices are surprisingly intrusive from the animals’ point of view as seen by the images of their reactions to them. On the other hand, although my motives have been “tested” a couple of times by coyotes (I’ve always walked away when this happens), this is rare. Although they’ve watched me, they effectively have not reacted to me, a human photographing them: this is why I’m able to document their natural family lives. We have plenty of other humans passing through the parks in San Francisco so I just blend in with the rest of them. If these had been non-urban coyotes not used to having humans around, I’m sure I would not be able to do what I do.

Conversely, coyotes DO routinely react to trap cameras, often adversely, and they are as aware of them as they are of any human in their area.

For the coyote, I think it boils down to an “understandable known” which is a human taking photos (and I’m talking about doing so respectfully and from a respectable distance), versus an “incomprehensible unknown” which are the clicks and whirrs and flashes of a contraption without a human presence yet “triggered” and focused on them when they appear: that’s when the lights and the noises go on. I’ve even seen several coyotes test this: they stand in the distance waiting for the IR lamp to turn off or the noises to begin, and then they make a slight movement that confirms THEY are the triggers.

So how are these field cameras intrusive? Coyotes often become startled by them, stopping dead in their tracks. The cameras elicit stares and investigative or wary reactions, especially as the coyote gets closer to them. The cameras all make sounds, and coyotes can hear even the faintest of noises, even if you can’t. So even if these cameras are missed visually by coyotes, they still know they are there — this must make them even more disturbing. Even the “black glow” cameras, where the IR flash is invisible to humans, are very visible to coyotes. I’ve actually seen coyotes interact with these cameras by angrily defecating in front of them, kicking dirt in anger at them, and even taking them down when they can [see photos and video below]. Wow! 

When the field cameras are up high and unreachable, they appear to be ignored or not to be seen, or so I thought, until I looked closer. When I set them on high ledges where they could be accessed by animals — they were.  So it’s a fallacy that trap cameras are not intrusive: coyotes know these are human instruments and likely malevolent in some way. Coyotes may even know you are spying on them.

“In the name of science” is often accepted as a good excuse for using them. Scientists want their “facts”, but people should know that they do impact the animals on deeper levels than they might be aware of. 

Having said all this, I have used trap cameras, as I stated above. Recently, my cameras were left up overtime when I neglected to pick them up for several days because I was too busy to go get them, and I discovered that the cameras caught some behaviors worth posting, always sprinkled with the reactions I’ve just described.

Some of what the cameras caught include: Dad bringing back prey to his pups, a mom regurgitating her food for her only pup, a youngster waiting around impatiently for his parents to return from hunting, older sibling playing with a younger pup,  a mom playing with her youngsters as though she were one of them, parents trying to shoo their youngster into hiding in the bushes but youngster doesn’t heed them, a dad pacing and standing very still as he listens carefully for a possible intruder he’s heard! I’ll post some of these in the coming weeks. The video today shows a pup exploring with insatiable curiosity, and Dad patrolling before that.

Some images and a video of coyotes encountering trap cameras.  Here are examples of how coyotes reacted with suspicion & investigation, kicking dirt in anger, defecating in anger and as a message, and knocking down the camera [Press one of the images to scroll through the gallery].

Another example: https://www.ktvq.com/entertainment/trending/video-extra-we-said-no-cameras

The Need for Wildness, by Tara Lohan

Press the above image or this link to read on: https://therevelator.org/coyotes-san-francisco/

Habitat Destruction

Two Coyotes, A Deceased Owl, and A Pussy Cat

Coyote stories usually involve more than just what meets the eye: background details and previous situations can contribute to filling out an understanding of the story. Then again, all stories are simply slices of time — partial stories. We tell them with a satisfying or instructive outcome or conclusion, but the story, in some form, of course, goes on. This entire observation lasted only a few minutes, but it was interesting, especially when expanded upon with a few things I know about these particular coyotes. But you’ll have to use your imagination to fill in the gaps: it’s still only a partial story.

Two coyotes were out foraging for mice in their field, when one of them came upon a find-of-the-week treasure: a dead barn owl. Coyotes are opportunistic eaters, and here, an opportunity presented itself and was acted upon.

An important word, again, about RAT POISON because that’s probably what killed the owl. Rat poison, or rodenticide, has been found in every owl we’ve taken up to WildCare for a necropsy. The owl would have eaten poisoned rats and mice which killed him/her. In the necropsies we’ve had done, the rodenticide was repeatedly found laced throughout the entire critter’s body. So this is probably the condition of this found dead owl. And now, coyotes have it. The poison in one owl probably isn’t enough to kill a coyote, but it can affect them, slowing down their reaction times and dulling their nerves: this is what rodenticide does. For instance, their chances of being hit by a car become greater. Cars are primary coyote killers in urban areas. And another tidbit about scavenging coyotes: they clean up carrion (dead animals) which keeps diseases from spreading — the dead owl at this point was carrion.

To continue. The female coyote grabbed the owl and ran with it to keep it away from the 2nd coyote, with the 2nd coyote at her heels. These two coyotes tease each other with mice which they grab from each other when the other coyote is unaware, but also I’ve seen that when the female finds prey on her own, she’ll keep and bury it just for herself so the other coyote won’t be able to find it. In these ways, coyotes interact with each other and food: teasing, sharing, not sharing.

internet photo

I ran to catch up with the coyotes but missed any more shots for this story. What I saw was the coyote with the owl clenched in her mouth running away from the second coyote as they zig-zagged their way around a community garden, and then, interestingly, a large orange tabby cat, right at their tails, zig-zagging along after them. Hmmm. Another tidbit of information about these two coyotes: they are scared of cats and run from them.

I lost track of them all until the two coyotes emerged with kind-of victory grins, but no owl. Had they buried it, or had the cat claimed it by scaring the coyotes away? I tend to think the lead coyote buried the owl to hide it as I saw her do a couple of days earlier with a road-killed raccoon she had found. But who would she be hiding it from? You can be sure that the interested animals following her know exactly where she cached it.

Both coyotes then climbed the hill above where they had been zig-zagging along. The cat was gone. The second coyote, the one who had followed the one with the owl, went off to hunt a little. But the first coyote plopped herself down within view of where the owl would have been buried and kept her eye on that area — until two dogs appeared and chased her away from her lookout post. But the dogs had no idea what she had been guarding, or even that she had been guarding anything at all. Dogs simply like to pursue coyotes.

My story was going to end there, leaving readers to guess who ultimately got the prized owl, but two days later I found the smelly old owl carcass pretty much still intact, but far from where the two coyotes and cat were seen with it. The coyotes may have simply been using the carcass as a toy, teasing and playing “keep-away” from each other. I wondered why it hadn’t been eaten, and I wondered if coyotes can sense rat poison and that the bird had been ill. I don’t know the answer, but since no coyotes were around, I took the opportunity to bag the carcass and dispose of it into a trash bin, to keep our coyotes safe from the high possibility of rat poison. The time had passed when a necropsy could have been accurately performed.

Scout Update: Heartwarming and Heartbreaking

I’ve been able to keep track of our territorially-displaced coyote, Scout, for the last four months as she kept herself out of the way and inconspicuous for the most part in various neighborhoods throughout the center of the city. And then, during the last several weeks she appeared in none of those places and I lost track of her.

So it was overwhelmingly heartwarming to hear from my friends that she had been spotted in her old territory the morning of June 26th: she’s alive and kicking and has not, apparently, given up hope of returning to her old home: what a trooper!  And a brave one at that since she might have been risking her life!  I was sent images, and indeed it was her: she was there, off-and-on, during a five hour interval that morning where, I’m told, she could be seen repeatedly yawning, I’m sure out of anxiety. I myself was only able to catch trail cam images of her before dawn that day as she headed toward her old hill (see above), and then again as she headed away that evening.

Wired making sure Scout stays away

By that evening, Wired — the radio-collared coyote who won the territory in a battle in February — had picked up her scent and was already on her trail. When Scout sensed her presence she was compelled to leave the area. The next day I caught images of her at one of her nearby hideouts where she hadn’t been in almost a month. Then, Wired, too, was seen checking out that same area (see images directly below): she is still hotly pursuing Scout. And now, again, I have lost track of Scout. Wired is back and remains in charge of what is now HER hill where she can be spotted sporadically.

The infrared photos show that Scout has several new sores on her underside and legs which she did not have before. Are these due to malnutrition, bugs, infections, being repelled by other coyotes, or simply wear and tear from life? Let’s hope this is just part and parcel of normal coyote nomad existence. Coyotes are very resilient, as this coyote has already shown us, so we can hope whatever is going on resolves itself quickly.

So Wired remains the territorial empress of what had belonged to Scout, and Scout remains the territorial-less roving nomad and vagrant, eeking out a living in-between territories claimed by other coyotes who keep her at bay. That is the part that’s heartrending.

Wired: the reigning alpha coyote

Please LEASH UP: Coyotes are entitled to defend their den areas here in San Francisco


This video from a field camera covers five hours. It was taken two months ago. I’ve cut out most of the non-action spaces, except those between a dog’s entrance into the area and the coyote’s coming out to “message” that dog to get out of her area. First thing to notice is that the coyote is a mother who is lactating — notice her underside. She needs dogs to stay away. A coyote is entitled to defend herself and her den area — her only tool with which to do so is her teeth. This is “defensive” behavior — it is not “aggressive” — this is not an “aggressive” coyote.

Please listen to the video: You’ll hear one owner brag about her dog always going where she tells it to — hmmm. You’ll hear a short scuffle and then the startled and freaked-out shriek of a dog — most likely the result of seeing the coyote’s snarly face and receiving a messaging leg-pinch, but the coyote may have gone further and actually nipped the dog. You’ll hear a man scornfully yell at the coyote to “get outa here”: this coyote is simply trying to keep dogs away from where her pups are hidden.  Remember, all you need to do when you see a coyote is to walk away from it with your dog leashed — you may have to resort to dragging your dog behind you as you walk away.

Almost all dogs are interested in the smells. This is one of the reasons they need to be kept leashed during pupping season: they should not be investigating den sites or near-den sites: it’s intrusive and stressful for coyote parents and potentially deadly for any pups. It also sets up the dog for a nip.

Please let’s give coyote parents some peace for raising their youngsters at the same time that we keep our dogs safe: all you need to do is leash-up and walk away from them, and keep your dogs from exploring off the beaten paths! The second coyote who came out was a father.

By the way, a couple of parents have allowed their children to crawl into such openings in our more naturally-wild parks. Maybe the openings look like they could become exciting little “forts” in the woods. Indeed that’s what they are — they’re already taken and belong to the wild critters who live there. There are plenty of signs everywhere throughout our parks advising that there are coyotes around. Please understand that coyotes NEED to protect their newborn pups. IF a child is nipped, there will be some tears and possibly a small wound to the child, but also it would be a tragedy because the coyote herself would most likely be euthanized — for simply protecting her pups. In the more overgrown woodsy parts of the parks we also have raccoons, skunks and plenty of rats who you should stay away from.

All About Coyote Pups: A Primer

I’ve put together some information on coyote pups based predominantly on my own observations. Other information is included, including some hearsay, which I’ve stated as such. Push the underlined links I’ve provided for even more information.

Coyote pups are about two months old now here in San Francisco.

Etiquette: Please stay away from any place where you even suspect a den might be, and please especially keep your dogs away from those areas. Besides cars, dogs are coyotes’ biggest threat in urban areas. You can prevent stress and trauma to yourself, your dog, and coyote parents with new pups by leashing your dogs if you must move through such an area. Word-of-mouth from other park visitors about a den’s general whereabouts should be enough for you to voluntarily avoid those areas.

Terminology: Baby coyotes are referred to as “pups” (not cubs)!

Dates of birth: In San Francisco, pups are normally born towards the beginning of April after a gestation of 63 days. Coyotes come into heat just once a year (as opposed to dogs’  twice a year cycle) — they can only produce one litter a year which is usually born sometime in the Springtime. The courting behavior which leads to pups can be observed at the end of January.

Litter Sizes: A “litter” is the group of pups that are born and raised together in one family. Some yearlings from the previous year’s litter or before — those that didn’t disperse (leave) — may remain in the family and may even help out a little by bringing in food for the youngsters. I have seen anywhere from one to seven pups born in one litter in San Francisco: that’s quite a spread!  I have heard of larger litters, as many as 11 to 19 (not in San Francisco), but I’ve also heard (but not seen) that this very large litter size may actually be the result of “den sharing”: two females sharing the same den.

From my own observations, I’ve seen litters vary in size over several consecutive years from the same parents as follows: Maeve and Toughy 1-2;  Ma’am and Monte 2-5-1; Cai and Yote 7-2-? (haven’t had a chance to count this year’s yet); Maya and Ivan (3-4-5); Chert and Silver 1-0-0-0-4;. These are all variations of normal litter sizes. The exception was the five litters where there were sometimes no survivals: this anomaly was almost certainly related to inbreeding in this family; Chert went through the motions of having pups during those three years of no pups, but none ever appeared. Besides these normal, small fluctuations, the litter sizes I’ve seen in San Francisco have not changed over the 12 years that I’ve been studying coyotes, even though the coyote population here has grown from an initial 8 in 2002 to several score and appears to be at its saturation point, as revealed by more and more brutal territorial battles that have displaced resident coyotes.

So although I’ve heard it speculated that coyotes “regulate” their litter size: I’ve not seen an indication of this here in San Francisco. They appear to just ‘get what they get’ — like the rest of us (those who don’t take hormones). And rather than the proviso that “locations dictate litter size”,  I would think that litter size might have a genetic component. I know that natural multiple births seem to run in human families, so might litter size also have a genetic component?

Newborn pup size: Coyote pups at birth weigh between 1/2 and 1 pound. This has been measured by wildlife rehabilitation facilities which take in these newborns. Weight can vary substantially between the pups in one family which may include some “runts”. Their personalities also develop differently. Some, right from the start are adventuresome and exploratory: they are curious about everything and wander far, rather brazenly. Others have more of a built-in wariness and fear: they are on-edge and more anxious and keep closer to home. Some are more interactive and social, involved in a lot of rough-house playing and teasing, while others are more withdrawn, preferring to sit back and watch the others. Each of these has it’s benefits for survival: one is not necessarily a “better” personality type than another.

Dens and birthing: Coyote pups are born in dens which were either dug for this purpose by their parents or they were pre-existing “homes” of other animals, such as raccoons or skunks, that were broadened and enlarged by the to-be coyote parents for their purposes.  A den can also be an existing hole in the ground, cavities under tree trunks, a depression with good covering, or a nook under a ledge. I have only seen dens long after they were abandoned, I have never seen an occupied den: I keep my distance from these.

Coyote dens are used only for a short time: for birthing and nursing during the very early life of pups and for the few months thereafter. Dens are outgrown, just as are birds’ nests or human cradles, after which the youngsters sleep in protected or hidden areas as do the parents. At around six weeks of age pups begin venturing beyond the den, but they’ll stick close to the den for much longer than that.

Here is a video of a wolf giving birth: I think we can assume that it must be pretty similar to what a coyote goes through. They emit howls as they give birth, and you hear a little of that here:

Do mothers stay in the dens for an extended period of time after giving birth? Well, if some do [this video shows pups right after, and pups’ first howls], it’s not a hard-and-fast behavior practice:  I’ve seen one mother out within a day of giving birth and I’ve seen others out in far less than a week.

Coyote pups may be left all alone for extended periods of time while the parents go off hunting. So, please don’t “save” what you think are “abandoned” pups — parents are likely simply off hunting. NO ONE can raise coyote pups as well as their own parents. You are actually hurting their chances for survival if you take their welfare into your own hands. If you truly believe they’ve been abandoned, then monitor them for several days before taking them to a wildlife center.

Nourishment: Coyote pups live on their mother’s milk alone for the first few weeks. This diet is soon supplemented with regurgitated food at about one month of age. The regurgitated food consists of whatever the parents consume, mostly rodents, but also fruit, amphibians, insects, birds and even garbage. How long do the youngsters lactate? I’ve seen mothers’ tit size shrink by the end of May, so I assume by that time the youngsters are well accustomed to semi-solid diets and on their way to whole prey soon. First, dead prey is brought to the youngsters, after which the pups are taken out to practice hunting on their own. Interestingly, I’ve seen coyotes over one year of age still being brought regurgitated food from a father: the youngsters approach Dad from below his chin, wedging their muzzles under his upper lip. This causes the regurgitation response. Dad then expels the food onto the ground, and the youngsters grab and eat it.

As the youngsters grow, Dad continues his contributions in the food department by bringing home prey to help feed the youngsters. Both coyote parents, unusually, raise the youngsters and you may find Dad at times sitting out and “minding” the offspring, giving Mom a break.

An interesting observation I’ve made is what I call “facilitating.” When the pups are learning to hunt, a mother will kill a rodent and  bury it where the pups can easily find it!

Survival rate:  Survival rate of pups is known to be low — 20-30% is what the literature says, but what I’ve seen in San Francisco is closer to about 70% from the time the pups can first be seen. However, it should be noted that I myself never know how many pups were actually born because I never intrude into their den areas: I wait to see what a litter is like until Mom brings them out or they are spotted through the bushes. Between birth and when I see them, there could have been more pups who didn’t make it.

First appearances: You might glimpse youngsters now, at two months of age, through the bushes, but it won’t be because coyotes want you to. Youngsters are highly protected as they grow up. And if a youngster sees you, he/she is bound to head straight to the bushes to hide! And parents are bound to come out to warn you to stay far, far away!

Family howl session in response to sirens. The howling family consists of Mom and Dad and four two-month old pups! Note that, although coyotes keep youngsters hidden as much as possible, they seem to have no qualms about being heard!

Here is what coyotes look like during their first year of growth:

Coyotes may leave home (dispersal) anytime after they are about 9 months of age, which is about January in SF. The earliest I’ve seen was a female who left home at 9 months of age. And the latest I’ve seen was a male who didn’t leave until he was 2.5 years of age, even though his father attempted pushing him out almost a year beforehand. The Presidio has tracked some of these dispersals to 60 miles south of the city, and I’ve been able to track some to vacated niches within the city itself. Many dispersing youngsters get killed by cars during this treacherous dispersal time in their lives. They also come across extremely hostile territorial coyotes who make it clear that they are not one bit welcome.

Keeping Their Secrets As Best They Can


Coyotes’ best kept secret is their pups — or at least they try to keep them a secret. If you happen to see any, it will be because the youngsters ventured beyond where their parents’ were trying so hard to keep them well hidden. The pups I know are all about two months old now here in San Francisco, as seen in the video.

And here is this family’s response to sirens: Mom, Dad, and FOUR youngsters:  Although they might not want to be seen, they don’t seem to mind being heard!!

Please stay away from any place where you even suspect a den might be, and please — specifically — keep your dogs away from those areas. It’s so easy to prevent stress and trauma to coyote families with new pups by leashing your dogs and keeping away. Word-of-mouth from other park visitors about the existence of pups should be enough for you to voluntarily avoid those areas with your unleashed dog. Parents will protect and defend their youngsters: why not just avoid the situation to begin with?

If a very young pup notices that s/he is being observed, s/he will run for cover.

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